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  • Forest and Grassland 

    forest is an ecosystem characterized by a dense community of trees.[2] Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function.[3][4][5] The United Nations’ Food and Agriculture Organization (FAO) defines a forest as, “Land spanning more than 0.5 hectares with trees higher than 5 meters and a canopy cover of more than 10 percent, or trees able to reach these thresholds in situ. It does not include land that is predominantly under agricultural or urban use.”[6] Using this definition, Global Forest Resources Assessment 2020 found that forests covered 4.06 billion hectares (10.0 billion acres; 40.6 million square kilometres; 15.7 million square miles), or approximately 31 percent of the world’s land area in 2020.[7]

    Forests are the largest terrestrial ecosystems of Earth by area, and are found around the globe.[8] 45 percent of forest land is in the tropical latitudes. The next largest share of forests are found in subarctic climates, followed by temperate, and subtropical zones.[9]

    Forests account for 75% of the gross primary production of the Earth’s biosphere, and contain 80% of the Earth’s plant biomassNet primary production is estimated at 21.9 gigatonnes of biomass per year for tropical forests, 8.1 for temperate forests, and 2.6 for boreal forests.[8]

    Forests form distinctly different biomes at different latitudes and elevations, and with different precipitation and evapotranspiration rates.[10] These biomes include boreal forests in subarctic climates, tropical moist forests and tropical dry forests around the Equator, and temperate forests at the middle latitudes. Forests form in areas of the Earth with high rainfall, while drier conditions produce a transition to savanna. However, in areas with intermediate rainfall levels, forest transitions to savanna rapidly when the percentage of land that is covered by trees drops below 40 to 45 percent.[11] Research conducted in the Amazon rainforest shows that trees can alter rainfall rates across a region, releasing water from their leaves in anticipation of seasonal rains to trigger the wet season early. Because of this, seasonal rainfall in the Amazon begins two to three months earlier than the climate would otherwise allow.[12][13] Deforestation in the Amazon and anthropogenic climate change hold the potential to interfere with this process, causing the forest to pass a threshold where it transitions into savanna.[14]

    Deforestation threatens many forest ecosystems. Deforestation occurs when humans remove trees from a forested area by cutting or burning, either to harvest timber or to make way for farming. Most deforestation today occurs in tropical forests. The vast majority of this deforestation is because of the production of four commodities: woodbeefsoy, and palm oil.[15] Over the past 2,000 years, the area of land covered by forest in Europe has been reduced from 80% to 34%. Large areas of forest have also been cleared in China and in the eastern United States,[16] in which only 0.1% of land was left undisturbed.[17] Almost half of Earth’s forest area (49 percent) is relatively intact, while 9 percent is found in fragments with little or no connectivity. Tropical rainforests and boreal coniferous forests are the least fragmented, whereas subtropical dry forests and temperate oceanic forests are among the most fragmented. Roughly 80 percent of the world’s forest area is found in patches larger than 1 million hectares (2.5 million acres). The remaining 20 percent is located in more than 34 million patches around the world – the vast majority less than 1,000 hectares (2,500 acres) in size.[9]

    Human society and forests can affect one another positively or negatively.[18] Forests provide ecosystem services to humans and serve as tourist attractions. Forests can also affect people’s health. Human activities, including unsustainable use of forest resources, can negatively affect forest ecosystems.[19]

    Definitions

    Forest in the Scottish Highlands

    Although the word forest is commonly used, there is no universally recognised precise definition, with more than 800 definitions of forest used around the world.[5] Although a forest is usually defined by the presence of trees, under many definitions an area completely lacking trees may still be considered a forest if it grew trees in the past, will grow trees in the future,[20] or was legally designated as a forest regardless of vegetation type.[21][22]

    There are three broad categories of definitions of forest in use: administrative, land use, and land cover.[21] Administrative definitions are legal designations, and may not reflect the type of vegetation that grows upon the land; an area can be legally designated “forest” even if no trees grow on it.[21] Land-use definitions are based on the primary purpose the land is used for. Under a land-use definition, any area used primarily for harvesting timber, including areas that have been cleared by harvesting, disease, fire, or for the construction of roads and infrastructure, are still defined as forests, even if they contain no trees. Land-cover definitions define forests based upon the density of trees, area of tree canopy cover, or area of the land occupied by the cross-section of tree trunks (basal area) meeting a particular threshold.[21] This type of definition depends upon the presence of trees sufficient to meet the threshold, or at least of immature trees that are expected to meet the threshold once they mature.[21]

    Under land-cover definitions, there is considerable variation on where the cutoff points are between a forest, woodland, and savanna. Under some definitions, to be considered a forest requires very high levels of tree canopy cover, from 60% to 100%,[23] which excludes woodlands and savannas, which have a lower canopy cover. Other definitions consider savannas to be a type of forest, and include all areas with tree canopies over 10%.[20]

    Some areas covered with trees are legally defined as agricultural areas, for example Norway spruce plantations, under Austrian forest law, when the trees are being grown as Christmas trees and are below a certain height.

    Etymology

    Since the 13th century, the Niepołomice Forest in Poland has had special use and protection. In this view from space, different coloration can indicate different functions.[24]

    The word forest derives from the Old French forest (also forès), denoting “forest, vast expanse covered by trees”; forest was first introduced into English as the word denoting wild land set aside for hunting[25] without necessarily having trees on the land.[26] Possibly a borrowing, probably via Frankish or Old High German, of the Medieval Latin foresta, denoting “open wood”, Carolingian scribes first used foresta in the capitularies of Charlemagne, specifically to denote the royal hunting grounds of the king. The word was not endemic to the Romance languages, e.g., native words for forest in the Romance languages derived from the Latin silva, which denoted “forest” and “wood(land)” (cf. the English sylva and sylvan; the Italian, Spanish, and Portuguese selva; the Romanian silvă; the Old French selve). Cognates of forest in Romance languages—e.g., the Italian foresta, Spanish and Portuguese floresta, etc.—are all ultimately derivations of the French word.

    A forest near VinitsaNorth Macedonia

    The precise origin of Medieval Latin foresta is obscure. Some authorities claim the word derives from the Late Latin phrase forestam silvam, denoting “the outer wood”; others claim the word is a Latinisation of the Frankish *forhist, denoting “forest, wooded country”, and was assimilated to forestam silvam, pursuant to the common practice of Frankish scribes. The Old High German forst denoting “forest”; Middle Low German vorst denoting “forest”; Old English fyrhþ denoting “forest, woodland, game preserve, hunting ground” (English frith); and Old Norse fýri, denoting “coniferous forest“; all of which derive from the Proto-Germanic *furhísa-, *furhíþija-, denoting “a fir-woodconiferous forest”, from the Proto-Indo-European *perkwu-, denoting “a coniferous or mountain forest, wooded height” all attest to the Frankish *forhist.

    Uses of forest in English to denote any uninhabited and unenclosed area are presently considered archaic.[27] The Norman rulers of England introduced the word as a legal term, as seen in Latin texts such as Magna Carta, to denote uncultivated land that was legally designated for hunting by feudal nobility (see royal forest).[27][28]

    These hunting forests did not necessarily contain any trees. Because that often included significant areas of woodland, “forest” eventually came to connote woodland in general, regardless of tree density.[citation needed] By the beginning of the fourteenth century, English texts used the word in all three of its senses: common, legal, and archaic.[27] Other English words used to denote “an area with a high density of trees” are firthfrithholtwealdwoldwood, and woodland. Unlike forest, these are all derived from Old English and were not borrowed from another language. Some present classifications reserve woodland for denoting a locale with more open space between trees, and distinguish kinds of woodlands as open forests and closed forests, premised on their crown covers.[29] Finally, sylva (plural sylvae or, less classically, sylvas) is a peculiar English spelling of the Latin silva, denoting a “woodland”, and has precedent in English, including its plural forms. While its use as a synonym of forest, and as a Latinate word denoting a woodland, may be admitted; in a specific technical sense it is restricted to denoting the species of trees that comprise the woodlands of a region, as in its sense in the subject of silviculture.[30] The resorting to sylva in English indicates more precisely the denotation that the use of forest intends.

    Evolutionary history

    The first known forests on Earth arose in the Middle Devonian (approximately 390 million years ago), with the evolution of cladoxylopsid plants like Calamophyton.[31] Appeared in the Late DevonianArchaeopteris was both tree-like and fern-like plant, growing to 20 metres (66 ft) in height or more.[32] It quickly spread throughout the world, from the equator to subpolar latitudes.[32] It is the first species known to cast shade due to its fronds and forming soil from its roots. Archaeopteris was deciduous, dropping its fronds onto the forest floor, the shade, soil, and forest duff from the dropped fronds creating the early forest.[32] The shed organic matter altered the freshwater environment, slowing its flow and providing food. This promoted freshwater fish.[32]

    Ecology

    Main article: Forest ecology

    Temperate rainforest in Tasmania’s Hellyer Gorge

    Forests account for 75% of the gross primary productivity of the Earth’s biosphere, and contain 80% of the Earth’s plant biomass.[8] Biomass per unit area is high compared to other vegetation communities. Much of this biomass occurs below ground in the root systems and as partially decomposed plant detritus. The woody component of a forest contains lignin, which is relatively slow to decompose compared with other organic materials such as cellulose or carbohydrate. The world’s forests contain about 606 gigatonnes of living biomass (above- and below-ground) and 59 gigatonnes of dead wood. The total biomass has decreased slightly since 1990, but biomass per unit area has increased.[33]

    Forest ecosystems broadly differ based on climate; latitudes 10° north and south of the equator are mostly covered in tropical rainforest, and the latitudes between 53°N and 67°N have boreal forest. As a general rule, forests dominated by angiosperms (broadleaf forests) are more species-rich than those dominated by gymnosperms (conifermontane, or needleleaf forests), although exceptions exist. The trees that form the principal structural and defining component of a forest may be of a great variety of species (as in tropical rainforests and temperate deciduous forests), or relatively few species over large areas (e.g., taiga and arid montane coniferous forests). The biodiversity of forests also encompasses shrubs, herbaceous plants, mossesfernslichensfungi, and a variety of animals.

    Trees rising up to 35 meters (115 ft) in height add a vertical dimension to the area of land that can support plant and animal species, opening up numerous ecological niches for arboreal animal species, epiphytes, and various species that thrive under the regulated microclimate created under the canopy.[34] Forests have intricate three-dimensional structures that increase in complexity with lower levels of disturbance and greater variety of tree species.[35]

    The biodiversity of forests varies considerably according to factors such as forest type, geography, climate, and soils – in addition to human use.[36] Most forest habitats in temperate regions support relatively few animal and plant species, and species that tend to have large geographical distributions, while the montane forests of Africa, South America, Southeast Asia, and lowland forests of Australia, coastal Brazil, the Caribbean islands, Central America, and insular Southeast Asia have many species with small geographical distributions.[36] Areas with dense human populations and intense agricultural land use, such as Europe, parts of Bangladesh, China, India, and North America, are less intact in terms of their biodiversity.[36] Northern Africa, southern Australia, coastal Brazil, Madagascar, and South Africa are also identified as areas with striking losses in biodiversity intactness.[36]

    Components

    Even, dense old-growth stand of beech trees (Fagus sylvatica) prepared to be regenerated by their saplings in the understory, in the Brussels part of the Sonian Forest.

    A forest consists of many components that can be broadly divided into two categories: biotic (living) and abiotic (non-living). The living parts include treesshrubsvinesgrasses and other herbaceous (non-woody) plants, mossesalgaefungiinsectsmammalsbirdsreptilesamphibians, and microorganisms living on the plants and animals and in the soil, connected by mycorrhizal networks.[37]

    Layers

    Spiny forest at Ifaty, Madagascar, featuring various Adansonia (baobab) species, Alluaudia procera (Madagascar ocotillo) and other vegetation

    The main layers of all forest types are the forest floor, the understory, and the canopy. The emergent layer, above the canopy, exists in tropical rainforests. Each layer has a different set of plants and animals, depending upon the availability of sunlight, moisture, and food.

    • The Forest floor is covered in dead plant material such as fallen leaves and decomposing logs, which detritivores break down into new soil. The layer of decaying leaves that covers the soil is necessary for many insects to overwinter and for amphibians, birds, and other animals to shelter and forage for food. Leaf litter also keeps the soil moist, stops erosion, and protects roots against extreme heat and cold.[38] The fungal mycelium that helps form the mycorrhizal network transmits nutrients from decaying material to trees and other plants. The forest floor supports a variety of plants, ferns, grasses, and tree seedlings, as well as animals such as antsamphibiansspiders, and millipedes.
    • Understory is made up of bushes, shrubs, and young trees that are adapted to living in the shade of the canopy.
    • Canopy is formed by the mass of intertwined branches, twigs, and leaves of mature trees. The crowns of the dominant trees receive most of the sunlight. This is the most productive part of the trees, where maximum food is produced. The canopy forms a shady, protective “umbrella” over the rest of the forest.
    • Emergent layer exists in a tropical rain forest and is composed of a few scattered trees that tower over the canopy.[39]

    In botany and countries like Germany and Poland, a different classification of forest vegetation is often used: tree, shrub, herb, and moss layers (see stratification (vegetation)).

    Types

    Casentinesi Forests, Italy

    Forests are classified differently and to different degrees of specificity. One such classification is in terms of the biomes in which they exist, combined with leaf longevity of the dominant species (whether they are evergreen or deciduous). Another distinction is whether the forests are composed predominantly of broadleaf trees, coniferous (needle-leaved) trees, or mixed.

    The number of trees in the world, according to a 2015 estimate, is 3 trillion, of which 1.4 trillion are in the tropics or sub-tropics, 0.6 trillion in the temperate zones, and 0.7 trillion in the coniferous boreal forests. The 2015 estimate is about eight times higher than previous estimates, and is based on tree densities measured on over 400,000 plots. It remains subject to a wide margin of error, not least because the samples are mainly from Europe and North America.[40]

    Forests can also be classified according to the amount of human alteration. Old-growth forest contains mainly natural patterns of biodiversity in established seral patterns, and they contain mainly species native to the region and habitat. In contrast, secondary forest is forest regrowing following timber harvest and may contain species originally from other regions or habitats.[41]

    Different global forest classification systems have been proposed, but none has gained universal acceptance.[42] UNEPWCMC‘s forest category classification system is a simplification of other, more complex systems (e.g. UNESCO‘s forest and woodland ‘subformations’). This system divides the world’s forests into 26 major types, which reflect climatic zones as well as the principal types of trees. These 26 major types can be reclassified into 6 broader categories: temperate needleleaf, temperate broadleaf and mixed, tropical moist, tropical dry, sparse trees and parkland, and forest plantations.[42] Each category is described in a separate section below.

    Temperate needleleaf

    Temperate needleleaf forests mostly occupy the higher latitudes of the Northern Hemisphere, as well as some warm temperate areas, especially on nutrient-poor or otherwise unfavourable soils. These forests are composed entirely, or nearly so, of coniferous species (Coniferophyta). In the Northern Hemisphere, pines Pinusspruces Picealarches Larixfirs Abies, Douglas firs Pseudotsuga, and hemlocks Tsuga make up the canopy; but other taxa are also important. In the Southern Hemisphere, most coniferous trees (members of Araucariaceae and Podocarpaceae) occur mixed with broadleaf species, and are classed as broadleaf-and-mixed forests.[42]

    Temperate broadleaf and mixed

    Broadleaf forest in Bhutan

    Temperate broadleaf and mixed forests include a substantial component of trees of the Anthophyta group. They are generally characteristic of the warmer temperate latitudes, but extend to cool temperate ones, particularly in the southern hemisphere. They include such forest types as the mixed deciduous forests of the United States and their counterparts in China and Japan; the broadleaf evergreen rainforests of Japan, Chile, and Tasmania; the sclerophyllous forests of Australia, central Chile, the Mediterranean, and California; and the southern beech Nothofagus forests of Chile and New Zealand.[42]

    Tropical moist

    There are many different types of tropical moist forests, with lowland evergreen broad-leaf tropical rainforests: for example várzea and igapó forests and the terra firme forests of the Amazon Basin; the peat swamp forestsdipterocarp forests of Southeast Asia; and the high forests of the Congo BasinSeasonal tropical forests, perhaps the best description for the colloquial term “jungle“, typically range from the rainforest zone 10 degrees north or south of the equator, to the Tropic of Cancer and Tropic of Capricorn. Forests located on mountains are also included in this category, divided largely into upper and lower montane formations, on the basis of the variation of physiognomy corresponding to changes in altitude.[43]

    Tropical dry

    Tropical dry forests are characteristic of areas in the tropics affected by seasonal drought. The seasonality of rainfall is usually reflected in the deciduousness of the forest canopy, with most trees being leafless for several months of the year. Under some conditions, such as less fertile soils or less predictable drought regimes, the proportion of evergreen species increases and the forests are characterised as “sclerophyllous“. Thorn forest, a dense forest of low stature with a high frequency of thorny or spiny species, is found where drought is prolonged, and especially where grazing animals are plentiful. On very poor soils, and especially where fire or herbivory are recurrent phenomena, savannas develop.[42]

    Sparse trees and savanna

    Sparse trees and savanna are forests with sparse tree-canopy cover. They occur principally in areas of transition from forested to non-forested landscapes. The two major zones in which these ecosystems occur are in the boreal region and in the seasonally dry tropics. At high latitudes, north of the main zone of boreal forestland, growing conditions are not adequate to maintain a continuously closed forest cover, so tree cover is both sparse and discontinuous. This vegetation is variously called open taiga, open lichen woodland, and forest tundra. A savanna is a mixed woodlandgrassland ecosystem characterized by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to support an unbroken herbaceous layer that consists primarily of grasses. Savannas maintain an open canopy despite a high tree density.[42]

    Plantations

    Forest plantations are generally intended for the production of timber and pulpwood. Commonly mono-specific, planted with even spacing between the trees, and intensively managed, these forests are generally important as habitat for native biodiversity. Some are managed in ways that enhance their biodiversity protection functions and can provide ecosystem services such as nutrient capital maintenance, watershed and soil structure protection and carbon storage.[41][42]

    Area

    Share of land that is covered by forest

    The annual net loss of forest area has decreased since 1990, but the world is not on track to meet the target of the United Nations Strategic Plan for Forests to increase forest area by 3 percent by 2030.[36]

    Share of forest area in total land area, top countries (2021)

    While deforestation is taking place in some areas, new forests are being established through natural expansion or deliberate efforts in other areas. As a result, the net loss of forest area is less than the rate of deforestation; and it, too, is decreasing: from 7.8 million hectares (19 million acres) per year in the 1990s to 4.7 million hectares (12 million acres) per year during 2010–2020.[36] In absolute terms, the global forest area decreased by 178 million hectares (440 million acres; 1,780,000 square kilometres; 690,000 square miles) between 1990 and 2020, which is an area about the size of Libya.[36]

    Societal significance

    Main articles: ForestryLogging, and Deforestation

    Redwood tree in northern California redwood forest, where many redwood trees are managed for preservation and longevity, rather than being harvested for wood production
    Burned forest on Thasos

    Ecosystem services

    Forests provide a diversity of ecosystem services including:

    • Converting carbon dioxide into oxygen and biomass. A full-grown tree produces about 100 kilograms (220 lb) of net oxygen per year.[44]
    • Acting as a carbon sink. Therefore, they are necessary to mitigate climate change.[45]
    • Aiding in regulating climate. For example, research from 2017 shows that forests induce rainfall. If the forest is cut, it can lead to drought,[46] and in the tropics to occupational heat stress of outdoor workers.[47]
    • Purifying water.
    • Mitigating natural hazards such as floods.
    • Serving as a genetic reserve.
    • Serving as a source of lumber and as recreational areas.
    • Serving as a source of woodlands and trees for millions of people dependent almost entirely on forests for subsistence for their essential fuelwood, food, and fodder needs.[48]

    The main ecosystem services can be summarized in the next table:[49]

    Type of forestCarbon storedBiodiversityOther
    Primary Boreal Forests1,042 billion tonnes of carbon, more than currently found in the atmosphere, 2 times more than all human caused emissions since the year 1870.Biodiversity services given by Canada forest alone are estimated as 703 billion dollars per year. Important for almost half of the birds in North America.Contain 60% of world surface freshwater.
    Primary Temperate Forests119 billion tonnes (like all CO2 emitted by humans in 2005–2017)Old growth forest has very high biodiversity. Some species link terrestrial ecosystems to marine.Some trees can live 1,000 years providing many services to humans. Help to protect people from floods and droughts.
    Primary Tropical Forests471 billion tonnes (more than all CO2 emissions from fossil fuel industry from the year 1750)Contain about two thirds of all species of terrestrial animals and plants.Creates clouds, rainfall.

    Some researchers state that forests do not only provide benefits, but can in certain cases also incur costs to humans.[50][51] Forests may impose an economic burden,[52][53] diminish the enjoyment of natural areas,[54] reduce the food-producing capacity of grazing land[55] and cultivated land,[56] reduce biodiversity,[57][58] reduce available water for humans and wildlife,[59][60] harbour dangerous or destructive wildlife,[50][61] and act as reservoirs of human and livestock disease.[62][63]

    An important consideration regarding carbon sequestration is that forests can turn from a carbon sink to a carbon source if plant diversity, density or forest area decreases, as has been observed in different tropical forests[64][65][66] The typical tropical forest may become a carbon source by the 2060s.[67] An assessment of European forests found early signs of carbon sink saturation, after decades of increasing strength.[68] The Intergovernmental Panel on Climate Change (IPCC) concluded that a combination of measures aimed at increasing forest carbon stocks, andsustainable timber offtake will generate the largest carbon sequestration benefit.[69]

    Forest-dependent people

    The term forest-dependent people is used to describe any of a wide variety of livelihoods that are dependent on access to forests, products harvested from forests, or ecosystem services provided by forests, including those of Indigenous peoples dependent on forests.[70] In India, approximately 22 percent of the population belongs to forest-dependent communities, which live in close proximity to forests and practice agroforestry as a principal part of their livelihood.[71] People of Ghana who rely on timber and bushmeat harvested from forests and Indigenous peoples of the Amazon rainforest are also examples of forest-dependent people.[70] Though forest-dependence by more common definitions is statistically associated with poverty and rural livelihoods, elements of forest-dependence exist in communities with a wide range of characteristics. Generally, richer households derive more cash value from forest resources, whereas among poorer households, forest resources are more important for home consumption and increase community resilience.[72]

    Indigenous peoples

    Forests are fundamental to the culture and livelihood of indigenous people groups that live in and depend on forests,[73] many of which have been removed from and denied access to the lands on which they lived as part of global colonialism. Indigenous lands contain 36% or more of intact forest worldwide, host more biodiversity, and experience less deforestation.[74][75][76] Indigenous activists have argued that degradation of forests and indigenous peoples’ marginalization and land dispossession are interconnected.[77][78] Other concerns among indigenous peoples include lack of Indigenous involvement in forest management and loss of knowledge related for the forest ecosystem.[79] Since 2002, the amount of land that is legally owned by or designated for indigenous peoples has broadly increased, but land acquisition in lower-income countries by multinational corporations, often with little or no consultation of indigenous peoples, has also increased.[80] Research in the Amazon rainforest suggests that indigenous methods of agroforestry form reservoirs of biodiversity.[81] In the U.S. state of Wisconsin, forests managed by indigenous people have more plant diversity, fewer invasive species, higher tree regeneration rates, and higher volume of trees.[82]

    Management

    Main article: Forest management

    World production of selected forest products

    Forest management has changed considerably over the last few centuries, with rapid changes from the 1980s onward, culminating in a practice now referred to as sustainable forest management. Forest ecologists concentrate on forest patterns and processes, usually with the aim of elucidating cause-and-effect relationships. Foresters who practice sustainable forest management focus on the integration of ecological, social, and economic values, often in consultation with local communities and other stakeholders.

    Priest River winds through mountains with a checkerboard design of trees to its east
    Priest River winding through Whitetail Butte with lots of forestry to the east—these lot patterns have existed since the mid-19th century. The white patches reflect areas with younger, smaller trees, where winter snow cover shows up brightly to the astronauts. Dark green-brown squares are parcels

    Humans have generally decreased the amount of forest worldwide. Anthropogenic factors that can affect forests include logging, urban sprawl, human-caused forest firesacid raininvasive species, and the slash and burn practices of swidden agriculture or shifting cultivation. The loss and re-growth of forests lead to a distinction between two broad types of forest: primary or old-growth forest and secondary forest. There are also many natural factors that can cause changes in forests over time, including forest firesinsectsdiseases, weather, competition between species, etc. In 1997, the World Resources Institute recorded that only 20% of the world’s original forests remained in large intact tracts of undisturbed forest.[83] More than 75% of these intact forests lie in three countries: the boreal forests of Russia and Canada, and the rainforest of Brazil.

    According to Food and Agriculture Organization‘s (FAO) Global Forest Resources Assessment 2020, an estimated 420 million hectares (1.0 billion acres) of forest have been lost worldwide through deforestation since 1990, but the rate of forest loss has declined substantially. In the most recent five-year period (2015–2020), the annual rate of deforestation was estimated at 10 million hectares (25 million acres), down from 12 million hectares (30 million acres) annually in 2010–2015.[33]

    The forest transition

    The transition of a region from forest loss to net gain in forested land is referred to as the forest transition. This change occurs through a few main pathways, including increase in commercial tree plantations, adoption of agroforestry techniques by small farmers, or spontaneous regeneration when former agricultural land is abandoned. It can be motivated by the economic benefits of forests, the ecosystem services forests provide, or cultural changes where people increasingly appreciate forests for their spiritual, aesthetic, or otherwise intrinsic value.[84] According to the Special Report on Global Warming of 1.5 °C of the Intergovernmental Panel on Climate Change, to avoid temperature rise by more than 1.5 degrees above pre-industrial levels, there will need to be an increase in global forest cover equal to the land area of Canada (10 million square kilometres (3.9 million square miles)) by 2050.[45]

    China instituted a ban on logging, beginning in 1998, due to the erosion and flooding that it caused.[85] In addition, ambitious tree-planting programmes in countries such as China, India, the United States, and Vietnam – combined with natural expansion of forests in some regions – have added more than 7 million hectares (17 million acres) of new forests annually. As a result, the net loss of forest area was reduced to 5.2 million hectares (13 million acres) per year between 2000 and 2010, down from 8.3 million hectares (21 million acres) annually in the 1990s. In 2015, a study for Nature Climate Change showed that the trend has recently been reversed, leading to an “overall gain” in global biomass and forests. This gain is due especially to reforestation in China and Russia.[86] New forests are not equivalent to old growth forests in terms of species diversity, resilience, and carbon capture. On 7 September 2015, the FAO released a new study stating that over the last 25 years the global deforestation rate has decreased by 50% due to improved management of forests and greater government protection.[87][88]

    Proportion of forest in protected areas, by region, 2020[33]

    There is an estimated 726 million hectares (1.79 billion acres) of forest in protected areas worldwide. Of the six major world regions, South America has the highest share of forests in protected areas, at 31 percent. The area of such areas globally has increased by 191 million hectares (470 million acres) since 1990, but the rate of annual increase slowed in 2010–2020.[33]

    Smaller areas of woodland in cities may be managed as urban forestry, sometimes within public parks. These are often created for human benefits; Attention Restoration Theory argues that spending time in nature reduces stress and improves health, while forest schools and kindergartens help young people to develop social as well as scientific skills in forests. These typically need to be close to where the children live.

    Canada

    Main article: Forests of Canada

    Garibaldi Provincial ParkBritish Columbia

    Canada has about 4 million square kilometres (1.5 million square miles) of forest land. More than 90% of forest land is publicly owned and about 50% of the total forest area is allocated for harvesting. These allocated areas are managed using the principles of sustainable forest management, which include extensive consultation with local stakeholders. About eight percent of Canada’s forest is legally protected from resource development.[89][90] Much more forest land—about 40 percent of the total forest land base—is subject to varying degrees of protection through processes such as integrated land use planning or defined management areas, such as certified forests.[90]

    By December 2006, over 1.2 million square kilometres (460,000 square miles) of forest land in Canada (about half the global total) had been certified as being sustainably managed.[91] Clearcutting, first used in the latter half of the 20th century, is less expensive, but devastating to the environment; and companies are required by law to ensure that harvested areas are adequately regenerated. Most Canadian provinces have regulations limiting the size of new clear-cuts, although some older ones grew to 110 square kilometres (42 sq mi) over several years.

    The Canadian Forest Service is the government department which looks after Forests in Canada.

    Latvia

    Latvian Pine Forest in Ķegums Municipality

    Latvia has about 3.27 million hectares (8.1 million acres; 12,600 square miles) of forest land, which equates to about 50.5% of Latvia’s total area of 64,590 square kilometres (24,938 sq mi) 1.51 million hectares (3.7 million acres) of forest land (46% of total forest land) is publicly owned and 1.75 million hectares (4.3 million acres) of forest land (54% of the total) is in private hands. Latvia’s forests have been steadily increasing over the years, which is in contrast to many other nations, mostly due to the forestation of land not used for agriculture. In 1935, there were only 1.757 million hectares (4.34 million acres) of forest; today this has increased by more than 150%. Birch is the most common tree at 28.2%, followed by pine (26.9%), spruce (18.3%), grey alder (9.7%), aspen (8.0%), black alder (5.7%), oak/ash (1.2%), with other hardwood trees making up the rest (2.0%).[92][93]

    United States

    In the United States, most forests have historically been affected by humans to some degree, though in recent years improved forestry practices have helped regulate or moderate large-scale impacts. The United States Forest Service estimated a net loss of about 2 million hectares (4.9 million acres) between 1997 and 2020; this estimate includes conversion of forest land to other uses, including urban and suburban development, as well as afforestation and natural reversion of abandoned crop and pasture land to forest. In many areas of the United States, the area of forest is stable or increasing, particularly in many northern states. The opposite problem from flooding has plagued national forests, with loggers complaining that a lack of thinning and proper forest management has resulted in large forest fires.[94][95]

  • Grassland 

    grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other herbs. Grasslands occur naturally on all continents except Antarctica and are found in most ecoregions of the Earth. Furthermore, grasslands are one of the largest biomes on Earth and dominate the landscape worldwide.[1] There are different types of grasslands: natural grasslands, semi-natural grasslands,[2] and agricultural grasslands.[1] They cover 31–69% of the Earth’s land area.[3][4]

    Definitions

    [edit]

    Coxilhas (hills covered by grasslands) in the PampasRio Grande do Sul state, Brazil

    Included among the variety of definitions for grasslands are:

    • “…any plant community, including harvested forages, in which grasses and/or legumes make up the dominant vegetation.”[1]
    • “…terrestrial ecosystems dominated by herbaceous and shrub vegetation, and maintained by fire, grazing, drought and/or freezing temperatures.” (Pilot Assessment of Global Ecosystems, 2000)[1]
    • “A region with sufficient average annual precipitation (25-75 cm) to support grass…” (Stiling, 1999)[1]

    Semi-natural grasslands are a very common subcategory of the grasslands biome.[5] These can be defined as:

    • Grassland existing as a result of human activity (mowing or livestock grazing), where environmental conditions and the species pool are maintained by natural processes.[6]

    They can also be described as the following:

    • “Semi-natural grasslands are one of the world’s most biodiverse habitats on a small spatial scales.”[7]
    • “Semi-natural grasslands belong to the most species rich ecosystems in the world.”[8]
    • “…have been formed over the course of centuries through extensive grazing and mowing.”[7]
    • “…without the use of pesticides or fertilisers in modern times.”[9]

    There are many different types of semi-natural grasslands, e.g. hay meadows.[9]

    Evolutionary history

    [edit]

    The graminoids are among the most versatile life forms. They became widespread toward the end of the Cretaceous period, and coprolites of fossilized dinosaur feces have been found containing phytoliths of a variety of grasses that include grasses that are related to modern rice and bamboo.

    The appearance of mountains in the western United States during the Miocene and Pliocene epochs, a period of some 25 million years, created a continental climate favourable to the evolution of grasslands.[10]

    Around 5 million years ago during the Late Miocene in the New World and the Pliocene in the Old World, the first true grasslands occurred. Existing forest biomes declined, and grasslands became much more widespread. It is known that grasslands have existed in Europe throughout the Pleistocene (the last 1.8 million years).[9] Following the Pleistocene ice ages (with their glacials and interglacials), grasslands expanded in the hotter, drier climates, and began to become the dominant land feature worldwide.[10] Since the grasslands have existed for over 1.8 million years, there is high variability. For example steppe-tundra dominated in Northern and Central Europe whereas a higher amount of xerothermic grasslands occurred in the Mediterranean area.[9] Within temperate Europe, the range of types is quite wide and also became unique due to the exchange of species and genetic material between different biomes.

    The semi-natural grasslands first appeared when humans started farming. So for the use of agriculture, forests got cleared in Europe. Ancient meadows and pastures were the parts that were suitable for cultivation. The semi-natural grasslands were formed from these areas.[9] However, there’s also evidence for the local persistence of natural grasslands in Europe, originally maintained by wild herbivores, throughout the pre-Neolithic Holocene.[11][12] The removal of the plants by the grazing animals and later the mowing farmers led to co-existence of other plant species around. In the following, the biodiversity of the plants evolve. Also, the species that already lived there adapted to the new conditions.[9]

    Most of the grassland areas have been turned to arable fields and disappeared again. The grasslands permanently became arable cropping fields due to the steady decrease in organic matter.[13] Nowadays, semi-natural grasslands are rather located in areas that are unsuitable for agricultural farming.[9]

    Ecology

    [edit]

    Biodiversity

    [edit]

    Grasslands dominated by unsown wild-plant communities (“unimproved grasslands”) can be called either natural or “semi-natural” habitat. Although their plant communities are natural, their maintenance depends upon anthropogenic activities such as grazing and cutting regimes. The semi-natural grasslands contain many species of wild plants, including grasses, sedges, rushes, and herbs; 25 plant-species per 100 square centimeters can be found.[9] A European record that was found on a meadow in Estonia described 76 species of plants in one square meter.[9] Chalk downlands in England can support over 40 species per square meter.

    Black rhino

    In many parts of the world, few examples have escaped agricultural improvement (fertilizing, weed killing, plowing, or re-seeding). For example, original North American prairie grasslands or lowland wildflower meadows in the UK are now rare and their associated wild flora equally threatened. Associated with the wild-plant diversity of the “unimproved” grasslands is usually a rich invertebrate fauna; there are also many species of birds that are grassland “specialists”, such as the snipe and the little bustard.[14] Owing to semi-natural grasslands being referred to as one of the most-species rich ecosystems in the world and essential habitat for many specialists, also including pollinators,[8] there are many approaches to conservation activities lately.

    Agriculturally improved grasslands, which dominate modern intensive agricultural landscapes, are usually poor in wild plant species due to the original diversity of plants having been destroyed by cultivation and by the use of fertilizers.

    Almost 90% of the European semi-natural grasslands do not exist anymore due to political and economic reasons. This loss took place during the 20th century.[7] The ones in Western and Central Europe have almost disappeared completely. There are a few left in Northern Europe.[7]

    Unfortunately, a large amount of red-listed species are specialists of semi-natural grasslands and are affected by the landscape change due to agriculture of the last century.[15]

    The original wild-plant communities having been replaced by sown monocultures of cultivated varieties of grasses and clovers, such as perennial ryegrass and white clover. In many parts of the world, “unimproved” grasslands are one of the most threatened types of habitat, and a target for acquisition by wildlife conservation groups or for special grants to landowners who are encouraged to manage them appropriately.

    Vegetation

    [edit]

    Quercus robur, also known as the English oak, dominating the semi-natural grasslands

    Grassland vegetation can vary considerably depending on the grassland type and on how strong it is affected by human impact. Dominant trees for the semi-natural grassland are Quercus roburBetula pendulaCorylus avellanaCrataegus and many kinds of herbs.[16]

    In chalk grassland, the plants can vary from very tall to very short. Quite tall grasses can be found in North American tallgrass prairieSouth American grasslands, and African savanna. Woody plants, shrubs or trees may occur on some grasslands—forming savannas, scrubby grassland or semi-wooded grassland, such as the African savannas or the Iberian deheza.[17]

    As flowering plants and trees, grasses grow in great concentrations in climates where annual rainfall ranges between 500 and 900 mm (20 and 35 in).[18] The root systems of perennial grasses and forbs form complex mats that hold the soil in place.

    Fauna

    [edit]

    Mountain plover

    Grasslands support the greatest aggregations of large animals on Earth, including jaguars, African wild dogs, pronghornblack-footed ferretplains bisonmountain plover, African elephant, Sunda tiger, black rhino, white rhino, savanna elephant, greater one-horned rhino, Indian elephant and swift fox. Grazing animals, herd animals, and predators in grasslands, like lions and cheetahs live in the grasslands of the African savanna.[19] Mites, insect larvaenematodes, and earthworms inhabit deep soil, which can reach 6 metres (20 feet) underground in undisturbed grasslands on the richest soils of the world. These invertebrates, along with symbiotic fungi, extend the root systems, break apart hard soil, enrich it with urea and other natural fertilizers, trap minerals and water and promote growth. Some types of fungi make the plants more resistant to insect and microbial attacks.[20]

    Cheetah

    Grassland in all its form supports a vast variety of mammals, reptiles, birds, and insects. Typical large mammals include the blue wildebeestAmerican bisongiant anteater, and Przewalski’s horse.[21]

    The plants and animals that live in grasslands are connected through an unlimited web of interactions. But the removal of key species—such as buffalo and prairie dogs within the American West—and introduction of invasive species, like cane toads in northern Australia, have disrupted the balance in these ecosystems and damaged a number of other species.[19] Grasslands are home to a number of the foremost magnificent animals on the planet—elephants, bison, lions—and hunters have found them to be enticing prey. But when hunting is not controlled or is conducted illegally, species can become extinct.[19]

    Ecosystem services

    [edit]

    Grasslands provide a range of marketed and non-marketed ecosystem services that are fundamental to the livelihoods of an estimated one billion people globally.[22]

    Carbon sequestration

    [edit]

    Grasslands hold about twenty percent of global soil carbon stocks.[3] Herbaceous (non-wooded) vegetation dominates grasslands and carbon is stored in the roots and soil underground. Above-ground biomass carbon is relatively short-lived due to grazing, fire, and senescence. Grassland species have an extensive fibrous root system, with grasses often accounting for 60-80% of the biomass carbon in this ecosystem. This underground biomass can extend several meters below the surface and store abundant carbon into the soil, resulting in deep, fertile soils with high organic matter content. For this reason, soil carbon accounts for about 81% of the total ecosystem carbon in grasslands. The close link between soil carbon and underground biomass leads to similar responses of these carbon pools to fluctuations in annual precipitation and temperature on a broad spatial scale. Because plant productivity is limited by grassland precipitation, carbon stocks are highest in regions where precipitation is heaviest, such as the high grass prairie in the humid temperate region of the United States. Similarly, as annual temperatures rise, grassland carbon stocks decrease due to increased evapotranspiration.[23]

    Grasslands have suffered large losses of organic carbon due to soil disturbances, vegetation degradation, fires, erosion, nutrient deficiencies, and water shortages. The type, frequency and intensity of the disturbance can play a key role in the soil organic carbon (SOC) balance of grasslands. Bedrock, irrigation practices, soil acidificationliming, and pasture management can all have potential impacts on grassland organic carbon stocks.[24] Good grassland management can reverse historical soil carbon losses.[3][25] The relationship of improved biodiversity with carbon storage is subject of research.[26]

    There is a lack of agreement on the amount of carbon that can be stored in grassland ecosystem. This is partly caused by different methodologies applied to measure soil organic carbon and limited respective datasets. Further, carbon accumulation in soils changes significantly over time and point in time measurements produce an insufficient evidence base.[27]

    Other ecosystem services

    [edit]

    • promotion of genetic diversity
    • weather amelioration[28]
    • provision of wildlife habitat

    Degradation

    [edit]

    Grasslands are among the most threatened ecosystems.[29] Global losses from grassland degradation are estimated to be over $7 billion per year.[30] According to the International Union for the Conservation of Nature (IUCN), the most significant threat to grasslands is human land use, especially agriculture and mining.[31] The vulnerability of grasslands stems from a range of factors, such as misclassification, poor protection and cultivation.[22]

    Causes

    [edit]

    Land use intensification

    [edit]

    Grasslands have an extensive history of human activity and disturbance.[32] To feed a growing human population, most of the world’s grasslands are converted from natural landscapes to fields of corn, wheat or other crops. Grasslands that have remained largely intact thus far, such as the East African savannas, are in danger of being lost to agriculture.[19] Grasslands are very sensitive to disturbances, such as people hunting and killing key species, or plowing the land to make more space for farms.

    Grassland vegetation is often a plagioclimax; it remains dominant in a particular area usually due to grazing, cutting, or natural or man-made fires, all discouraging colonization by and survival of tree and shrub seedlings.[33] Some of the world’s largest expanses of grassland are found in the African savanna, and these are maintained by wild herbivores as well as by nomadic pastoralists and their cattle, sheep or goats. Grasslands have an impact on climate change by slower decomposition rates of litter compared to forest environments.[34]

    Main land-cover trajectories from the 1960s to 2015

    Grasslands may occur naturally or as a result of human activity. Hunting cultures around the world often set regular fires to maintain and extend grasslands and prevent fire-intolerant trees and shrubs from taking hold. The tallgrass prairies in the U.S. Midwest may have been extended eastward into IllinoisIndiana, and Ohio by human agency. Much grassland in northwest Europe developed after the Neolithic Period when people gradually cleared the forest to create areas for raising their livestock.[35]

    Climate change

    [edit]

    Grasslands often occur in areas with annual precipitation is between 600 mm (24 in) and 1,500 mm (59 in) and average mean annual temperatures ranges from −5 and 20 °C.[36] However, some grasslands occur in colder (−20 °C) and hotter (30 °C) climatic conditions. Grassland can exist in habitats that are frequently disturbed by grazing or fire, as such disturbance prevents the encroachment of woody species. Species richness is particularly high in grasslands of low soil fertility such as serpentine barrens and calcareous grasslands, where woody encroachment is prevented as low nutrient levels in the soil may inhibit the growth of forest and shrub species. Another common predicament often experienced by the ill-fated grassland creatures is the constant burning of plants, fueled by oxygen and many expired photosynthesizing organisms, with the lack of rain pushing this problem to further heights.[37] When not limited by other factors, increasing CO2 concentration in the air increases plant growth, similarly as water use efficiency, which is very important in drier regions. However, the advantages of elevated CO2 are limited by factors including water availability and available nutrients, particularly nitrogen. Thus effects of elevated CO2 on plant growth will vary with local climate patterns, species adaptations to water limitations, and nitrogen availability. Studies indicate that nutrient depletion may happen faster in drier regions, and with factors like plant community composition and grazing. Nitrogen deposition from air pollutants and increased mineralization from higher temperatures can increase plant productivity, but increases are often among a discount in biodiversity as faster-growing plants outcompete others. A study of a California grassland found that global change may speed reductions in diversity and forb species are most prone to this process.[23]

    Afforestation or introduction of invasive species

    [edit]

    Misguided afforestation efforts, for example as part of the global effort to increase carbon sequestration, can harm grasslands and their core ecosystem services.[38][39] Forest centric restoration efforts can create the risk of misreading and misclassifying of landscapes.[22] A map created by the World Resources Institute in collaboration with the IUCN identifies 2 billion hectares for potential forest restoration. It is criticised for including 900 million hectares of grasslands.[40][41] It is expected that non-native grasses will continue to outperform native species under warmer and drier conditions that occur in many grasslands due to climate change.[42]

    Management

    [edit]

    The type of land management used in grasslands can also lead to grassland loss or degradation. Many grasslands and other open ecosystems depend on disturbances such as wildfirescontrolled burns or grazing to persist, although this subject is still controversial.[43] A study in Brazilian Subtropical Highland Grasslands found that grasslands without traditional land management—which uses fire every two years and extensive cattle grazing—can disappear within 30 years.[44] This study showed that grasslands inside protected areas, in which fire is not allowed and cattle grazing is banned, grasslands were quickly replaced by shrubs (shrub encroachment).

    Types of degradation

    [edit]

    Land cover change

    [edit]

    Land cover has always changed during the years. The following relates to the changes between 1960 and 2015. There has been a decrease in semi-natural grasslands and an increase in areas with arable land, forest and land used for infrastructure and buildings. The line style and relative thickness of the lines indicates the percentage of the total area that changed. Changes less than 1% and land-cover classes with all changes less than 1% (i.e. semi-natural wetlands and water) are not included.[15]

    In 1960 most of the land, 49.7%, was covered with forest and there was also more semi-natural grassland (18.8%) than arable land (15.8%). In 2015 this has changed drastically. The forest cover has increased (50.8%) and arable land has also increased (20.4%), but the semi-natural grassland cover has decreased. Although it still covers a large area of the earth (10.6%).[15]

    A quarter of semi-natural grassland was lost through intensification, i.e. it was converted into arable or pasture land and forests.[45] It is more likely that intensification will occur in flat semi-natural grasslands, especially if the soil is fertile. On the other hand, grasslands, where the land is drought-prone or less productive, are more likely to persist as semi-natural grasslands than grasslands with fertile soil and low gradient of the terrain.[46] Furthermore, the accessibility of the land is also important, as it is then easier to fertilize, for example. For instance, if it is located near a road. With the development of technology, it is becoming increasingly easy to cultivate land with a steeper gradient, to the detriment of grasslands. The management of grasslands is also changing permanently. There is increased use of mineral fertilizers, furthermore borders and field edges are removed to enlarge fields and leveling the terrain to facilitate the use of agricultural machinery.[15]

    The professional study of dry grasslands falls under the category of rangeland management, which focuses on ecosystem services associated with the grass-dominated arid and semi-arid rangelands of the world. Rangelands account for an estimated 70% of the earth’s landmass; thus, many cultures including those of the United States are indebted to the economics that the world’s grasslands have to offer, from producing grazing animals, tourism, ecosystems services such as clean water and air, and energy extraction.[47]

    Main article: Woody plant encroachment

    Vast areas of grassland are affected by woody encroachment, which is the expansion of woody plants at the expense of the herbaceous layer. Woody encroachment is caused by a combination of human impact (e.g. fire exclusion, overstocking and resulting overgrazing) and environmental factors (i.e. increased CO2 levels in the atmosphere). It can have severe negative consequences on key ecosystem services, like land productivity and groundwater recharge.

    Conservation and restoration

    [edit]

    Despite growing recognition of the importance of grasslands, understanding of restoration options remains limited.[48] Cost of grassland restoration is highly variable and respective data is scarce.[49] Successful grassland restoration has several dimensions, including recognition in policy, standardisation of indicators of degradation, scientific innovation, knowledge transfer and data sharing.[50]

    Restoration methods and measures include the following:[51]

    • prescribed fires
    • appropriate management of livestock and wild herbivores: in light of land use intensification caused by global food demand, grassland land use practices may need to be adjusted to better support key ecosystem services.[52]
    • tree cutting
    • shrub removal
    • invasive species control
    • reintroduction of native grasses and forbs via seeding or transplant: a main challenge for grassland restoration is how to overcome seed limitation.[48]

    For the period 2021–2030 the United Nations General Assembly has proclaimed the UN Decade on Restoration, involving a joint resolution by over 70 countries. It is led by the United Nations Environment Programme and the Food and Agriculture Organization.[53]

    Types of grasslands

    [edit]

    Meadow by the Desna river in Ukraine

    Classifications of grassland

    [edit]

    Grassland types by Schimper (1898, 1903):[54]

    • Meadow (hygrophilous or tropophilous grassland)
    • Steppe (xerophilous grassland)
    • Savannah (xerophilous grassland containing isolated trees)Steppe family: a common grassland animal, the swift fox

    Grassland types by Ellenberg and Mueller-Dombois (1967):[55]

    Formation-class V. Terrestrial herbaceous communities

    1. Savannas and related grasslands (tropical or subtropical grasslands and parklands)
    2. Steppes and related grasslands (e.g. North American “prairies” etc.)
    3. Meadows, pastures or related grasslands
    4. Sedge swamps and flushes
    5. Herbaceous and half-woody salt swamps
    6. Forb vegetationA hike through the Tallgrass Prairie Heritage Park in Canada

    Grassland types by Laycock (1979):[56]

    1. Tallgrass (true) prairie
    2. Shortgrass prairie
    3. Mixed-grass prairie
    4. Shrub steppe
    5. Annual grassland
    6. Desert (arid) grassland
    7. High mountain grassland

    General grasslands types

    [edit]

    Tropical and subtropical

    [edit]

    These grasslands can be classified as the tropical and subtropical grasslands, savannas and shrublands biome. The rainfall level for that grassland type is between 90 and 150 centimeters per year. Grasses and scattered trees are common for that ecoregion, as well as large mammals, such as wildebeest (Connochaetes taurinus) and zebra (Equus zebra). Notable tropical and subtropical grasslands include the Llanos grasslands of South America.[57]

    Cumberland Plain Woodland, a grassy woodland that covers Western Sydney

    Temperate

    [edit]

    Mid-latitude grasslands, including the prairie and Pacific grasslands of North America, the Pampas of ArgentinaBrazil and Uruguay, calcareous downland, and the steppes of Europe. They are classified with temperate savannas and shrublands as the temperate grasslands, savannas, and shrublands biome. Temperate grasslands are the home to many large herbivores, such as bisongazelleszebrasrhinoceroses, and wild horsesCarnivores like lionswolvescheetahs and leopards are also found in temperate grasslands. Other animals of this region include deerprairie dogsmicejack rabbitsskunkscoyotessnakesfoxesowlsbadgers, blackbirds, grasshoppersmeadowlarkssparrowsquailshawks and hyenas.[58]

    Flooded

    [edit]

    Grasslands that are flooded seasonally or year-round, like the Everglades of Florida, the Pantanal of BrazilBolivia and Paraguay or the Esteros del Ibera in Argentina, are classified with flooded savannas as the flooded grasslands and savannas biome and occur mostly in the tropics and subtropics. The species that live in these grasslands are well adapted to the hydrologic regimes and soil conditions. The Everglades—the world’s largest rain-fed flooded grassland—is rich in 11,000 species of seed-bearing plants, 25 species of orchids, 300 bird species, and 150 fish species.

    Water-meadows are grasslands that are deliberately flooded for short periods.[59]

    Grassland in the Antelope ValleyCalifornia

    Montane

    [edit]

    High-altitude grasslands located on high mountain ranges around the world, like the Páramo of the Andes Mountains. They are part of the montane grasslands and shrublands biome and can be tropical, subtropical, and temperate. The plants and animals, that can be found in the tropical montane, are able to adapt to cool, wet conditions as well as intense sunlight.[60]

    Tundra grasslands

    [edit]

    Similar to montane grasslands, polar Arctic tundra can have grasses, but high soil moisture means that few tundras are grass-dominated today. However, during the Pleistocene glacial periods (commonly referred to as ice ages), a grassland known as steppe-tundra or mammoth steppe occupied large areas of the Northern Hemisphere. These areas were very cold and arid and featured sub-surface permafrost (hence tundra) but were nevertheless productive grassland ecosystems supporting a wide variety of fauna. As the temperature increased and the climate became wetter at the beginning of the Holocene much of the mammoth steppe transitioned to forest, while the drier parts in central Eurasia remained as a grassland, becoming the modern Eurasian steppe.[61]

    Desert and xeric

    [edit]

    Also called desert grasslands, they are composed of sparse grassland ecoregions located in the deserts and xeric shrublands biome. Temperature extremes and low amounts of rainfall characterise these kinds of grasslands. Therefore, plants and animals are well adapted to minimize water loss.[62]

    Temperate grasslands, savannas, and shrublands ecoregions

    [edit]

    The grassland ecoregions of the temperate grasslands, savannas, and shrublands biome are:

    Afrotropical temperate grasslands, savannas, and shrublands ecoregionsvte
    Al Hajar montane woodlandsOmanUnited Arab Emirates
    Amsterdam and Saint-Paul Islands temperate grasslandsAmsterdam IslandSaint-Paul Island
    Tristan da Cunha–Gough Islands shrub and grasslandsTristan da CunhaGough Island
    Australasian temperate grasslands, savannas, and shrublands ecoregionsvte
    Canterbury–Otago tussock grasslandsNew Zealand
    Eastern Australia mulga shrublandsAustralia
    Southeast Australia temperate savannaAustralia
    Nearctic temperate grasslands, savannas, and shrublands ecoregionsvte
    California Central Valley grasslandsUnited States
    Canadian aspen forests and parklandsCanadaUnited States
    Central and Southern mixed grasslandsUnited States
    Central forest–grasslands transitionUnited States
    Central tall grasslandsUnited States
    Columbia PlateauUnited States
    Edwards Plateau savannaUnited States
    Flint Hills tall grasslandsUnited States
    Montana valley and foothill grasslandsUnited States
    Nebraska Sand Hills mixed grasslandsUnited States
    Northern mixed grasslandsCanadaUnited States
    Northern short grasslandsCanadaUnited States
    Northern tall grasslandsCanadaUnited States
    Palouse grasslandsUnited States
    Texas blackland prairiesUnited States
    Western short grasslandsUnited States
    Neotropical temperate grasslands, savannas, and shrublands ecoregionsvte
    Argentine EspinalArgentina
    Argentine MonteArgentina
    Humid PampasArgentinaUruguay
    Patagonian grasslandsArgentinaChileUnited Kingdom (Falkland Islands)
    Patagonian steppeArgentinaChileUnited Kingdom (Falkland Islands)
    Semi-arid PampasArgentina
    Palearctic temperate grasslands, savannas, and shrublands ecoregionsvte
    Alai–Western Tian Shan steppeKazakhstanTajikistanUzbekistan
    Altai steppe and semi-desertKazakhstan
    Central Anatolian steppeTurkey
    Daurian forest steppeChinaMongoliaRussia
    Eastern Anatolian montane steppeArmeniaAzerbaijanGeorgiaIranTurkey
    Emin Valley steppeChinaKazakhstan
    Faroe Islands boreal grasslandsFaroe IslandsDenmark
    Gissaro–Alai open woodlandsKyrgyzstanTajikistanUzbekistan
    Kazakh forest steppeKazakhstanRussia
    Kazakh steppeKazakhstanRussia
    Kazakh UplandsKazakhstan
    Mongolian–Manchurian grasslandChinaMongoliaRussia
    Pontic steppeKazakhstanMoldovaRomaniaRussiaUkraineBulgaria
    Sayan Intermontane steppeRussia
    Selenge–Orkhon forest steppeMongoliaRussia
    South Siberian forest steppeRussia
    Syrian xeric grasslands and shrublandsIraqJordanSyria
    Tian Shan foothill arid steppeChinaKazakhstanKyrgyzstan

    Tropical and subtropical grasslands, savannas, and shrublands ecoregions

    [edit]

    Afrotropical tropical and subtropical grasslands, savannas, and shrublands ecoregionsvte
    Angolan miombo woodlandsAngola
    Angolan mopane woodlandsAngolaNamibia
    Ascension scrub and grasslandsAscension Island
    Central Zambezian miombo woodlandsAngolaBurundiDemocratic Republic of the CongoMalawiTanzaniaZambia
    East Sudanian savannaCameroonCentral African RepublicChadDemocratic Republic of the CongoEritreaEthiopiaSouth SudanSudanUganda
    Eastern miombo woodlandsMozambiqueTanzania
    Guinean forest–savanna mosaicBeninBurkina FasoCameroonGambiaGhanaGuineaGuinea BissauIvory CoastNigeriaSenegalTogo
    Itigi–Sumbu thicketTanzaniaZambia
    Kalahari Acacia-Baikiaea woodlandsBotswanaNamibiaSouth AfricaZimbabwe
    Mandara Plateau mosaicCameroonNigeria
    Northern Acacia–Commiphora bushlands and thicketsEthiopiaKenyaSouth SudanUganda
    Northern Congolian forest–savanna mosaicCameroonCentral African RepublicDemocratic Republic of the CongoSouth SudanUganda
    Sahelian Acacia savannaBurkina FasoCameroonCape VerdeChadEritreaEthiopiaMaliMauritaniaNigerNigeriaSenegalSouth SudanSudan
    Serengeti volcanic grasslandsKenyaTanzania
    Somali Acacia–Commiphora bushlands and thicketsEritreaEthiopiaKenyaSomalia
    South Arabian fog woodlands, shrublands, and duneOmanSaudi ArabiaYemen
    Southern Acacia–Commiphora bushlands and thicketsKenyaTanzania
    Southern Africa bushveldBotswanaSouth AfricaZimbabwe
    Southern Congolian forest–savanna mosaicAngolaDemocratic Republic of the Congo
    Southern miombo woodlandsMalawiMozambiqueZambiaZimbabwe
    Saint Helena scrub and woodlandsSaint Helena
    Victoria Basin forest–savanna mosaicBurundiDemocratic Republic of the CongoEthiopiaKenyaRwandaSouth SudanTanzaniaUganda
    West Sudanian savannaBeninBurkina FasoGambiaGhanaGuinea,MaliIvory CoastNigerNigeriaSenegal
    Western Congolian forest–savanna mosaicAngolaDemocratic Republic of the CongoRepublic of the Congo
    Western Zambezian grasslandsAngolaZambia
    Zambezian and mopane woodlandsBotswanaEswatiniMalawiMozambiqueNamibiaSouth AfricaZambiaZimbabwe
    Zambezian Baikiaea woodlandsAngolaBotswanaNamibiaZambiaZimbabwe
    Australasian tropical and subtropical grasslands, savannas, and shrublands ecoregionsvte
    Arnhem Land tropical savannaAustralia
    Brigalow tropical savannaAustralia
    Cape York Peninsula tropical savannaAustralia
    Carpentaria tropical savannaAustralia
    Einasleigh Uplands savannaAustralia
    Kimberley tropical savannaAustralia
    Mitchell grass downsAustralia
    Trans-Fly savanna and grasslandsIndonesiaPapua New Guinea
    Victoria Plains tropical savannaAustralia
    Indomalayan tropical and subtropical grasslands, savannas, and shrublands ecoregionsvte
    Terai–Duar savanna and grasslandsBhutanIndiaNepal
    Nearctic tropical and subtropical grasslands, savannas, and shrublands ecoregionsvte
    Western Gulf coastal grasslandsMexicoUnited States
    Neotropical tropical and subtropical grasslands, savannas, and shrublands ecoregionsvte
    Beni savannaBolivia
    Campos rupestresBrazil
    CerradoBoliviaBrazilParaguay
    Clipperton Island shrub and grasslandsFrance (Clipperton Island)
    Córdoba montane savannaArgentina
    Guianan savannaBrazilGuyanaVenezuela
    Humid ChacoArgentinaBrazilParaguayUruguay
    LlanosVenezuelaColombia
    Uruguayan savannaArgentinaBrazilUruguay
    Oceanian tropical and subtropical grasslands, savannas, and shrublands ecoregionsvte
    Hawaiian tropical high shrublandsHawaiʻi
    Hawaiian tropical low shrublandsHawaiʻi
    Northwestern Hawaii scrubHawaiʻiMidway Atoll